Motivation 1 ('organisms-as-agents thesis') "says that organisms really do exhibit some or even all of the attributes of agency". Motivation 2 ('organism-as-agents heuristic') "says that it can be heuristically useful to treat organisms as if they were agents for certain intellectual purposes".
It's interesting that both motivations appear to be about modelling organisms as agents, as opposed to any other level of organisation. This feels like it misses some of the most interesting insights we might get from biological agency, namely those around agents at different levels of organisation interacting - e.g: ants and ant colonies, cancer cells and multicellular organisms, or individual organisms and selection pressures (which could be treated as as-if agents at evolutionary timescales).
It's interesting that both motivations appear to be about modelling organisms as agents, as opposed to any other level of organisation. This feels like it misses some of the most interesting insights we might get from biological agency, namely those around agents at different levels of organisation interacting - e.g: ants and ant colonies, cancer cells and multicellular organisms, or individual organisms and selection pressures (which could be treated as as-if agents at evolutionary timescales).